This article will be permanently flagged as inappropriate and made unaccessible to everyone. Are you certain this article is inappropriate? Excessive Violence Sexual Content Political / Social
Email Address:
Article Id: WHEBN0001787105 Reproduction Date:
Animal sexual behaviour takes many different forms, even within the same species. Common mating or reproductively motivated systems include monogamy, polyandry, polygamy, and promiscuity. Other sexual behaviour may be reproductively motivated (e.g. sex apparently due to duress or coercion and situational sexual behaviour) or non-reproductively motivated (e.g. interspecific sexuality, sexual arousal from objects or places, sex with dead animals, homosexual sexual behaviour, bisexual sexual behaviour, and a range of other behaviours).
When animal sexual behaviour is reproductively motivated, it is often termed mating or copulation; for most non-human mammals, mating and copulation occur at estrus (the most fertile period in the mammalian female's reproductive cycle), which increases the chances of successful impregnation.[1][2] Some animal sexual behaviour involves competition, sometimes fighting, between multiple males. Females often select males for mating only if they appear strong and able to protect themselves. The male that wins a fight may also have the chance to mate with a larger number of females and will therefore pass on his genes to their offspring.[3]
Historically, it was believed that only humans and a small number of other species performed sexual acts other than for reproduction, and that animals' sexuality was instinctive and a simple "stimulus-response" behaviour. A range of species masturbate and may use objects as tools to help them do so.
In sociobiology and behavioural ecology, the term "mating system" is used to describe the ways in which animal societies are structured in relation to sexual behaviour. The mating system specifies which males mate with which females, and under what circumstances. There are four basic systems:
Monogamy occurs when one male mates with one female exclusively. A monogamous mating system is one in which individuals form long-lasting pairs and cooperate in raising offspring. These pairs may last for a lifetime, such as in pigeons,[6] or it may occasionally change from one mating season to another, such as in emperor penguins.[7] In contrast with tournament species, these pair-bonding species have lower levels of male aggression, competition and little sexual dimorphism. Zoologists and biologists now have evidence that monogamous pairs of animals are not always sexually exclusive. Many animals that form pairs to mate and raise offspring regularly engage in sexual activities with extra-pair partners.[8][9][10][11] This includes previous examples, such as swans. Sometimes, these extra-pair sexual activities lead to offspring. Genetic tests frequently show that some of the offspring raised by a monogamous pair come from the female mating with an extra-pair male partner.[9][12][13][14] These discoveries have led biologists to adopt new ways of talking about monogamy According to Ulrich Reichard (2003):
Social monogamy refers to a male and female's social living arrangement (e.g., shared use of a territory, behaviour indicative of a social pair, and/or proximity between a male and female) without inferring any sexual interactions or reproductive patterns. In humans, social monogamy takes the form of monogamous marriage. Sexual monogamy is defined as an exclusive sexual relationship between a female and a male based on observations of sexual interactions. Finally, the term genetic monogamy is used when DNA analyses can confirm that a female-male pair reproduce exclusively with each other. A combination of terms indicates examples where levels of relationships coincide, e.g., sociosexual and sociogenetic monogamy describe corresponding social and sexual, and social and genetic monogamous relationships, respectively.[15]
Whatever makes a pair of animals socially monogamous does not necessarily make them sexually or genetically monogamous. Social monogamy, sexual monogamy, and genetic monogamy can occur in different combinations.
Social monogamy is relatively rare in the animal kingdom. The actual incidence of social monogamy varies greatly across different branches of the evolutionary tree. Over 90% of avian species are socially monogamous.[10][16] This stands in contrast to mammals. Only 3% of mammalian species are socially monogamous, although up to 15% of primate species are.[10][16] Social monogamy has also been observed in reptiles, fish, and insects.
Sexual monogamy is also rare among animals. Many socially monogamous species engage in extra-pair copulations, making them sexually non-monogamous. For example, while over 90% of birds are socially monogamous, "on average, 30% or more of the baby birds in any nest [are] sired by someone other than the resident male."[17] Patricia Adair Gowaty has estimated that, out of 180 different species of socially monogamous songbirds, only 10% are sexually monogamous.[18]
The incidence of genetic monogamy, determined by DNA fingerprinting, varies widely across species. For a few rare species, the incidence of genetic monogamy is 100%, with all offspring genetically related to the socially monogamous pair. But genetic monogamy is strikingly low in other species. Barash and Lipton note:
The highest known frequency of extra-pair copulations are found among the fairy-wrens, lovely tropical creatures technically known as Malurus splendens and Malurus cyaneus. More than 65% of all fairy-wren chicks are fathered by males outside the supposed breeding group.[16]p. 12
Such low levels of genetic monogamy have surprised biologists and zoologists, forcing them to rethink the role of social monogamy in evolution. They can no longer assume social monogamy determines how genes are distributed in a species. The lower the rates of genetic monogamy among socially monogamous pairs, the less of a role social monogamy plays in determining how genes are distributed among offspring.
Polygyny occurs when one male gets exclusive mating rights with multiple females. In some species, notably those with harem-like structures, only one of a few males in a group of females will mate. Technically, polygyny in sociobiology and zoology is defined as a system in which a male has a relationship with more than one female, but the females are predominantly bonded to a single male. Should the active male be driven out, killed, or otherwise removed from the group, in a number of species the new male will ensure that breeding resources are not wasted on another male's young.[19] The new male may achieve this in many different ways, including:
Von Haartman specifically described the mating behaviour of the European pied flycatcher as successive polygyny.[20] Within this system, the males leave their home territory once their primary female lays her first egg. Males then create a second territory, presumably in order to attract a secondary female to breed. Even when they succeed at acquiring a second mate, the males typically return to the first female to exclusively provide for her and her offspring.[21]
Polygynous mating structures are estimated to occur in up to 90% of mammal species.[22] As polygyny is the most common form of polygamy among vertebrates (including humans, to some extent), it has been studied far more extensively than polyandry or polygynandry.
Polyandry occurs when one female gets exclusive mating rights with multiple males. In some species, such as redlip blennies, both polygyny and polyandry are observed.[23]
The males in some deep sea gonads, which release sperm in response to hormones in the female's bloodstream indicating egg release. This extreme sexual dimorphism ensures that, when the female is ready to spawn, she has a mate immediately available.[24] A single anglerfish female can "mate" with many males in this manner.
Polygynandry occurs when multiple males mate indiscriminately with multiple females. The numbers of males and females need not be equal, and in vertebrate species studied so far, there are usually fewer males. Two examples of systems in primates are promiscuous mating chimpanzees and bonobos. These species live in social groups consisting of several males and several females. Each female copulates with many males, and vice versa. In bonobos, the amount of promiscuity is particularly striking because bonobos use sex to alleviate social conflict as well as to reproduce. This mutual promiscuity is the approach most commonly used by spawning animals, and is perhaps the "original fish mating system."[4]:161 Common examples are forage fish, such as herrings, which form huge mating shoals in shallow water. The water becomes milky with sperm and the bottom is draped with millions of fertilized eggs.[4]:161
The term polygamy is an umbrella term used to refer generally to non-monogamous matings. As such, polygamous relationships can be polygynous, polyandrous or polygynandrous. In a small number of species, individuals can display either polygamous or monogamous behaviour depending on environmental conditions. An example is the social wasp Apoica flavissima.
A tournament species is one in which "mating tends to be highly polygamous and involves high levels of male-male aggression and competition."[25] Tournament behaviour often correlates with high levels of sexual dimorphism, examples of species including chimpanzees and baboons. Most polygamous species present high levels of tournament behaviour, with a notable exception being bonobos.
Female and male sexual behaviour differ in many species. Often, males are more active in initiating mating, and bear the more conspicuous sexual ornamentation like antlers and colourful plumage. This is a result of anisogamy, where sperm are smaller and much less costly (energetically) to produce than eggs. This difference in physiological cost means that males are more limited by the number of mates they can secure, while females are limited by the quality of genes of her mates, a phenomenon known as Bateman's principle.[26] Many females also have extra reproductive burdens in that parental care often falls mainly, or exclusively, on them. Thus, females are more limited in their potential reproductive success.[27] In species where males take on more of the reproductive costs, such as sea horses and jacanas, the role is reversed, and the females are larger, more aggressive and more brightly coloured than the males.
In hermaphroditic animals, the costs of parental care can be evenly distributed between the sexes, e.g. earthworms. In some species of planarians, sexual behaviour takes the form of penis fencing. In this form of copulation, the individual that first penetrates the other with the penis, forces the other to be female, thus carrying the majority of the cost of reproduction.[28] Post mating, banana slugs will some times gnaw off their partners penis as an act of sperm competition.[29] In the grey slug, the sharing of cost leads to a spectacular display, where the mates suspend themselves high above the ground from a slime thread, ensuring none of them can refrain from taking on the cost of egg-bearer.[30]
Many animal species have specific mating (or breeding) periods e.g. (seasonal breeding) so that offspring are born or hatch at an optimal time. In marine species with limited mobility and external fertilization like corals, sea urchins and clams, the timing of the common spawning is the only externally visible form of sexual behaviour. In areas with continuously high primary production, some species have a series of breeding seasons throughout the year. This is the case with most primates (who are primarily tropical and subtropical animals). Some animals (opportunistic breeders) breed dependent upon other conditions in their environment aside from time of year.
Mating seasons are often associated with changes to herd or group structure, and behavioural changes, including territorialism amongst individuals. These may be annual (e.g. wolves), biannual (e.g. dogs) or more frequently (e.g. horses). During these periods, females of most mammalian species are more mentally and physically receptive to sexual advances, a period scientifically described as estrous but commonly described as being "in season" or "in heat". Sexual behaviour may occur outside estrus,[31] and such acts as do occur are not necessarily harmful.[32]
Some mammals (e.g. domestic cats, rabbits and camilidae) are termed "induced ovulators". For these species, the female ovulates due to an external stimulus during, or just prior, to mating, rather than ovulating cyclically or spontaneously. Stimuli causing induced ovulation include the sexual behaviour of coitus, sperm and pheromones. Domestic cats have penile spines. Upon withdrawal of a cat's penis, the spines rake the walls of the female's vagina, which may cause ovulation.[33][34]
For many amphibians, an annual breeding cycle applies, typically regulated by ambient temperature, precipitation, availability of surface water and food supply. This breeding season is accentuated in temperate regions, in boreal climate the breeding season is typically concentrated to a few short days in the spring.
Like many coral reef dwellers, the clownfish spawn around the time of the full moon in the wild. In a group of clownfish, there is a strict dominance hierarchy. The largest and most aggressive female is found at the top. Only two clownfish, a male and a female, in a group reproduce through external fertilization. Clownfish are sequential hermaphrodites, meaning that they develop into males first, and when they mature, they become females. If the female clownfish is removed from the group, such as by death, one of the largest and most dominant males will become a female. The remaining males will move up a rank in the hierarchy.
Various [35] Vasopressin is also involved in the sexual behaviour of some animals.[36]
The mating system of prairie voles is monogamous; after mating, they form a lifelong bond. In contrast, montane voles have a polygamous mating style. When montane voles mate, they form no attachments and separate after copulation. Studies on the brains of these two species have found that it is two neurohormones and their respective receptors that are responsible for these differences in mating strategies. Male prairie voles release vasopressin after copulation prairie vole and attachment to the female then develops. Female prairie voles release oxytocin after reproducing and an attachment to her partner also develops. Neither male of female montane voles have such a high quantity of oxytocin and vasopressin when they mate. Even when injected with these neurohormones, the mating system does not change. In contrast, if prairie voles are injected with the neurohormones, they may form a lifelong attachment, even if they have not mated. The reason for this is that prairie voles have more oxytocin and vasopressin receptors than do montane voles, and are therefore far more receptive to the two neurohormones. It is not the quantity of the hormone that determines the mating system and bond-formation, rather, it is number of neurohormone receptors.[36]
Mother rats experience a postparum estrus which makes them highly motivated to mate. However, they also have a strong motivation to protect their newly born pups. As a consequence, the mother rat solicits males to the nest but simultaneously becomes aggressive towards them to protect her young. If the mother rat is given injections of an oxytocin receptor antagonist, they no longer experience these maternal motivations.[37]
Prolactin influences social bonding in rats.[37]
Oxytocin plays a similar role in non-human primates as it does in humans.
Grooming, sex, and cuddling frequencies correlate positively with levels of oxytocin. As the level of oxytocin increases so does sexual motivation. While oxytocin plays a major role in parent child relationships, it is also found to play a role in adult sexual relationships. Its secretion affects the nature of the relationship or if there will even be a relationship at all.
Studies have shown that oxytocin is higher in monkeys in lifelong monogamous relationships compared to monkeys which are single. Furthermore, the oxytocin levels of the couples correlate positively; when the oxytocin secretion of one increases the other one also increases. Higher levels of oxytocin are related to monkeys expressing more behaviours such as cuddling, grooming and sex, while lower levels of oxytocin reduce motivation for these activities.
Research on oxytocin's role in the animal brain suggests that it plays less of a role in behaviours of love and affection than previously believed. "When oxytocin was first discovered in 1909, it was thought mostly to influence a mother’s labor contractions and milk let-down. Then, in the 1990s, research with prairie voles found that giving them a dose of oxytocin resulted in the formation of a bond with their future mate (Azar, 40)." Oxytocin has since been treated by the media as the sole player in the "love and mating game" in mammals. This view, however, is proving to be false as, "most hormones don’t influence behaviour directly. Rather, they affect thinking and emotions in variable ways (Azar, 40)." There is much more involved in sexual behaviour in the mammalian animal than oxytocin and vasopressin can explain.[38][39][40][41][42][43][44]
It is often assumed that animals do not have sex for pleasure, or alternatively that humans, pigs (and perhaps dolphins and one or two species of primate) are the only species that do. This is sometimes stated as "animals mate only for reproduction". This view is considered a misconception by some scholars.[45] [35] and the notion that non-human animals experience emotions similar to humans is a contentious subject.[47][48][49][50]
A 2006 Danish Animal Ethics Council report[51] which examined current knowledge of animal sexuality in the context of legal queries concerning sexual acts by humans, has the following comments, primarily related to domestically common animals:
Even though the evolution-related purpose of mating can be said to be reproduction, it is not actually the creating of offspring which originally causes them to mate. It is probable that they mate because they are motivated for the actual copulation, and because this is connected with a positive experience. It is therefore reasonable to assume that there is some form of pleasure or satisfaction connected with the act. This assumption is confirmed by the behaviour of males, who in the case of many species are prepared to work to get access to female animals, especially if the female animal is in oestrus, and males who for breeding purposes are used to having sperm collected become very eager, when the equipment they associate with the collection is taken out.[52]
There is nothing in [53]
Koinophilia is the love of the "normal" or phenotypically common (from the Greek, koinos, meaning "the usual" or "common").[54] The term was introduced to scientific literature in 1990, and refers to the tendency of animals seeking a mate to prefer that mate not to have any unusual, peculiar or deviant features.[54] Similarly, animals preferentially choose mates with low fluctuating asymmetry. [55] However, animal sexual ornaments can evolve through runaway selection, which is driven by (usually female) selection for non-standard traits.[56]
The field of study of sexuality in non-human species was a long-standing taboo.[57] In the past, researchers sometimes failed to observe, mis-categorizing and mis-described sexual behaviour which did not meet their preconceptions. In earlier periods, bias tended to support what would now be described as conservative sexual mores. An example of overlooking behaviour relates to descriptions of giraffe mating:
When nine out of ten pairings occur between males, "[e]very male that sniffed a female was reported as sex, while dominance, competition or greetings."[57]
In the 21st century, liberal social or sexual views are often projected upon animal subjects of research. Popular discussions of bonobos are a frequently cited example. Current research frequently expresses views such as that of the Natural History Museum at the University of Oslo, which in 2006 held an exhibition on animal sexuality:
"Many researchers have described homosexuality as something altogether different from sex. They must realise that animals can have sex with who they will, when they will and without consideration to a researcher's ethical principles."[57]
Other animal activities may be misinterpreted due to the frequency and context in which animals perform the behaviour. For example, domestic ruminants display behaviours such as mounting and head-butting. This often occurs when the animals are establishing dominance relationships and are not necessarily sexually-motivated. Careful analysis must be made to interpret what animal motivations are being expressed by those behaviours.[58]
Alternate male strategies which allow small males to engage in cuckoldry can develop in species such as fish where spawning is dominated by large and aggressive males. Cuckoldry is a variant of polyandry, and can occur with sneak spawners. A sneak spawner is a male that rushes in to join the spawning rush of a spawning pair.[59] A spawning rush occurs when a fish makes a burst of speed, usually on a near vertical incline, releasing gametes at the apex, followed by a rapid return to the lake or sea floor or fish aggregation.[60] Sneaking males do not take part in courtship. In salmon and trout, for example, jack males are common. These are small silvery males that migrate upstream along with the standard, large, hook-nosed males and that spawn by sneaking into redds to release sperm simultaneously with a mated pair. This behaviour is an evolutionarily stable strategy for reproduction, because it is favoured by natural selection just like the "standard" strategy of large males.[61]
gonochorism, where each individual in a species is either male or female, and remains that way throughout their lives. Most fish are gonochorists, but hermaphroditism is known to occur in 14 families of teleost fishes.[62]
Usually hermaphrodites are sequential, meaning they can switch sex, usually from female to male (protogyny). This can happen if a dominant male is removed from a group of females. The largest female in the harem can switch sex over a few days and replace the dominant male.[62] This is found amongst coral reef fishes such as groupers, parrotfishes and wrasses. It is less common for a male to switch to a female (protandry).[4]:162 As an example, most wrasses are protogynous hermaphrodites within a haremic mating system.[63][64] Hermaphroditism allows for complex mating systems. Wrasses exhibit three different mating systems: polygynous, lek-like, and promiscuous mating systems.[65]
Sexual cannibalism is a behaviour in which a female animal kills and consumes the male before, during, or after copulation. Sexual cannibalism confers fitness advantages to both the male and female. Sexual cannibalism is common among insects, arachnids[66] and amphipods.[66] There is also evidence of sexual cannibalism in gastropods and copepods.[67]
Sex in a forceful or apparently coercive context has been documented in a variety of species. In some herbivorous herd species, or species where males and females are very different in size, the male dominates sexually by force and size.
Some species of birds appear to combine sexual intercourse with apparent violent assault; these include ducks,[68][69] and geese. Female white-fronted bee-eaters are subjected to forced copulations. When females emerge from their nest burrows, males sometimes force them to the ground and mate with them. Such forced copulations are made preferentially on females who are laying and who may therefore lay eggs fertilised by the male.[70]
In the Acilius genus of water beetles, an "evolutionary arms race" between the two sexes means that there is no courtship system for these beetles. Their mating behaviours include males holding females underwater till exhausted, and allowing only occasional access to the surface to breathe for up to six hours (to prevent them breeding with other males), and females which have a variety of body shapings (to prevent males from gaining a grip).[71]
It has been reported that young male elephants in South Africa sexually coerced and killed rhinoceroses.[72] This interpretation of the elephants' behaviour was disputed by one of the original study's authors, who said there was "nothing sexual about these attacks".[73]
Parthenogenesis is a form of asexual reproduction in which growth and development of embryos occur without fertilization. Technically, parthenogenesis it is not a behaviour, however, sexual behaviours may be involved.
Whip-tailed lizard females have the ability to reproduce through parthenogenesis and as such males are rare and sexual breeding non-standard. Females engage in "pseudocopulation"[74] to stimulate ovulation, with their behaviour following their hormonal cycles; during low levels of estrogen, these (female) lizards engage in "masculine" sexual roles. Those animals with currently high estrogen levels assume "feminine" sexual roles. Lizards that perform the courtship ritual have greater fecundity than those kept in isolation due to an increase in hormones triggered by the sexual behaviours. So, even though asexual whiptail lizards populations lack males, sexual stimuli still increase reproductive success. From an evolutionary standpoint these females are passing their full genetic code to all of their offspring rather than the 50% of genes that would be passed in sexual reproduction.
It is rare to find true parthenogenesis in fishes, where females produce female offspring with no input from males. All-female species include the Texas silverside, Menidia clarkhubbsi[75] as well as a complex of Mexican mollies.[4]:162
Parthenogenesis has been recorded in 70 vertebrate species[76] including hammerhead sharks,[77] blacktip sharks,[78] amphibians[79][80] and crayfish.[81][82]
Unisexuality occurs when a species is all-male or all-female. Unisexuality occurs in some fish species, and can take complex forms. Squalius alburnoides, a minnow found in several river basins in Portugal and Spain, appears to be an all-male species. The existence of this species illustrates the potential complexity of mating systems in fish. The species originated as a hybrid between two species, and is diploid, but not hermaphroditic. It can have triploid and tetraploid forms, including all-female forms that reproduce mainly through hybridogenesis.[83]
There is a range of behaviours that animals perform which appear to be sexually motivated but which can not result in reproduction. These include -
Seahorses, once considered to be monogamous species with pairs mating for life, were described in a 2007 study as "promiscuous, flighty, and more than a little bit gay".[109] Scientists at 15 aquaria studied 90 seahorses of three species. Of 3,168 sexual encounters, 37% were same-sex acts. Flirting was common (up to 25 potential partners a day of both sexes); only one species (the British spiny seahorse) included faithful representatives, and for these 5 of 17 were faithful, 12 were not. Bisexual behaviour was widespread and considered "both a great surprise and a shock", with big-bellied seahorses of both sexes not showing partner preference. 1,986 contacts were male-female, 836 were female-female and 346 were male-male.[109]
The bonobo, which has a matriarchal society, is a fully bisexual species — both males and females engage in sexual behaviour with the same and the opposite sex, with females being particularly noted for engaging in sexual behaviour with each other and at up to 75% of sexual activity being nonreproductive. Primatologist Frans de Waal believes that bonobos use sexual activity to resolve conflict between individuals.[110] Sexual activity occurs between almost all ages and sexes of bonobo societies.[111]
Male
Selv om det evolutionsmæssige formål med at parre sig kan siges at være reproduktion, er det ikke selve det, at dyrene får afkom, der i første omgang får dem til at parre sig. Det er til gengæld sandsynligt, at de parrer sig, fordi de er motiverede for selve parringsakten, og at denne er forbundet med en positiv oplevelse. Det er derfor rimeligt at antage, at der er en eller anden form for behag eller tilfredsstillelse forbundet med akten. Denne antagelse bekræftes af adfærden hos handyr, der for mange arters vedkommende er parate til at arbejde for at få adgang til hundyr, især hvis hundyret er i brunst, og handyr der i avlsøjemed er vant til at få tappet sæd – de viser stor ivrighed, når det udstyr, de forbinder med sædopsamlingen, tages frem.
Der er intet ved hunpattedyrenes anatomi eller fysiologi, der modsiger, at stimulation af kønsorganerne og parring skulle kunne være en positiv oplevelse – fx fungerer klitoris på samme måde som hos kvinder. Videnskabelige undersøgelser har vist, at reproduktionssuccesen forbedres ved stimulation af klitoris på bl.a. køer og hopper i forbindelse med insemination, fordi det forbedrer sædtransporten pga. sammentrækninger af de indre kønsdele. Dette gælder sandsynligvis også hundyr af andre dyrearter, og sammentrækninger i de indre kønsdele ses fx også under orgasme hos kvinder. Det er derfor rimeligt at antage, at det seksuelle samvær kan være forbundet med en positiv oplevelse for hundyrene.
Research into human evolution confirms that, in some cases, interspecies sexual activity may have been responsible for the evolution of new species (speciation). Analysis of animal genes found evidence that after humans had diverged from other apes, interspecies mating nonetheless occurred regularly enough to change certain genes in the new gene pool.[128] Researchers found that the X chromosomes of humans and chimps may have diverged around 1.2 million years after the other chromosomes. One possible explanation is that modern humans emerged from a hybrid of human and chimp populations.[129] A 2012 study questioned this explanation, concluding that "there is no strong reason to involve complicated factors in explaining the autosomal data".[130]
Many animals make plugs of mucus to seal the female's orifice after mating. Normally such plugs are secreted by the male, to block subsequent partners. In spiders the female can assist the process.[127] Spider sex is unusual in that males transfer their sperm to the female on small limbs called pedipalps. They use these to pick their sperm up from their genitals and insert it into the female's sexual orifice, rather than copulating directly.[127] On the 14 occasions a sexual plug was made, the female produced it without assistance from the male. On ten of these occasions the male's pedipalps then seemed to get stuck while he was transferring the sperm (which is rarely the case in other species of spider), and he had great difficulty freeing himself. In two of those ten instances, he was eaten as a result.[127]
Butterflies spend much time searching for mates. When the male spots a mate, he will fly closer and closer and release pheromones. He then performs a special courtship dance to attract the female. If the female appreciates the dancing she may join him. Then they join their bodies together end to end at their abdomens. Here, the male passes the sperm to the female's egg-laying tube which will soon be fertilized by the sperm.[126] The male often dies shortly after mating.
Corals can be both gonochoristic (unisexual) and hermaphroditic, each of which can reproduce sexually and asexually. Reproduction also allows corals to settle new areas. Corals predominantly reproduce sexually. 25% of hermatypic corals (stony corals) form single sex (gonochoristic) colonies, while the rest are hermaphroditic.[123] About 75% of all hermatypic corals "broadcast spawn" by releasing gametes—eggs and sperm—into the water to spread offspring. The gametes fuse during fertilization to form a microscopic larva called a planula, typically pink and elliptical in shape.[124] Synchronous spawning is very typical on the coral reef and often, even when multiple species are present, all corals spawn on the same night. This synchrony is essential so that male and female gametes can meet. Corals must rely on environmental cues, varying from species to species, to determine the proper time to release gametes into the water. The cues involve lunar changes, sunset time, and possibly chemical signalling.[123] Synchronous spawning may form hybrids and is perhaps involved in coral speciation.[125]
Penis fencing is a mating behaviour engaged in by certain species of flatworm, such as Pseudobiceros bedfordi. Species which engage in the practice are hermaphroditic, possessing both eggs and sperm-producing testes.[122] The species "fence" using two-headed dagger-like penises which are pointed, and white in color. One organism inseminates the other. The sperm is absorbed through pores in the skin, causing fertilization.
Invertebrates are often calcium carbonate or chitin, and is called a love dart. Each snail manoeuvres to get its genital pore in the best position, close to the other snail's body. Then, when the body of one snail touches the other snail's genital pore, it triggers the firing of the love dart.[119] After the snails have fired their darts, they copulate and exchange sperm as a separate part of the mating progression. The love darts are covered with a mucus that contains a hormone-like substance that facilitates the survival of the sperm.[120][121]
Mammals mate by vaginal copulation. To achieve this, the male usually mounts the female from behind. The female may exhibit lordosis in which she arches her back ventrally to facilitate entry of the penis. Other than humans, only bonobos mate in a face-to-face position.[116] Some camelids mate in a lying-down position.[117] In most ruminant species, a single pelvic thrust occurs during copulation.[118]
[115] The female
[113] The man was found not guilty after it was revealed at trial that the dolphin was known to tow bathers through the water by hooking his penis around them.[113]
Human, Endangered Species, IUCN Red List, Animal, Congo River
Speciation, Open access, Genus, Class (biology), Order (biology)
Evolutionary psychology, Evolutionary biology, Richard Dawkins, Evolution, Sociology
Sexual selection, Mammals, Pollination, Breast, Ethology
Ethology, Animal communication, Memory, Primates, Neuroethology
Richard Dawkins, Biology, Ethology, Animal cognition, Animal communication
Psychology, Neuroethology, Biology, Nobel Prize in Physiology or Medicine, Animal communication
Lgbt, LGBT rights in France, Sexual orientation, Gender identity, Transgender
Gender identity, Transgender, Third gender, Lgbt, Asexuality